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. 2020 Feb 19;29(4):893–904. doi: 10.1002/pro.3835

Table 1.

Pilotins and accessory molecules of the T2SS, T3SS, and T4PS

System Pilotin Accessory molecule Cognate secretin Bacterial species Main references
T2SS EpsS

EpsA

EpsB

EpsD Vibrio vulnificus

Howard et al. 15

Strozen et al.55

AspS GspD Escherichia coli (EPEC)

Dunstan et al.38

Yin et al.36

AspS GspD Vibrio cholerae Dunstan et al.38
OutS OutD Erwinia chrysanthemi Rehman et al.61
PulS PulD Klebsiella oxytoca

Tosi et al.33

Nickerson et al.34

ExeA

ExeB

ExeD Aeromonas hydrophila

Li and Howard62

Ast et al.63

Vanderlinde et al.64

T3SS MxiM MxiJ MxiD Shigella flexneri

Schuch and Maurelli65

Lario et al.56

Okon et al.66

ExsB PscC Pseudomonas aeruginosa

Izoré et al.57

Perdu et al.67

YscW YscC Yersinia enterocolitica

Burghout et al.68

Ross and Plano132

InvH InvG Salmonella enterica

Daefler and Russell69

Craig and Koronakis70

YsaP YsaC Yersinia enterocolitica Rau and Darwin71
T4aPS PilW PilP PilQ Neisseria meningitidis

Trindade et al.59

Carbonnelle et al.72

Golovanov et al.58

PilF PilQ Pseudomonas aeruginosa Koo et al.60
Tgl PilQ Myxococcus xanthus Nudleman et al.73
T4bPS BfpG BfpB Escherichia coli Lieberman et al.74
TcpQ TcpC Vibrio cholerae Chang et al.75
FimV PilQ Pseudomonas aeruginosa Wehbi et al.76
CpaE CpaC Caulobacter crescentus Viollier et al.77
TadD RcpA Aggregatibacter actinomycetemcomitans Clock et al.78