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. 2013 Mar 21;20(3):316–331. doi: 10.1016/j.chembiol.2013.02.006

Table 1.

Proteins that Interact with PHBs

Binding Partners PHB Biological Consequences References
Mitochondrial Proteins

m-AAA protease PHB1 regulation of membrane protein degradation Steglich et al., 1999
ATAD3 PHB1 mitochondrial protein synthesis He et al., 2012
nucleoids (mitochondrial DNA associated to nucleoproteins) PHB1/2 maintenance of the organization and copy number of the mtDNA Kasashima et al., 2008
dynamin-like GTPase OPA-1 PHB1 stabilization of the structure of mitochondria during their fusion and morphogenesis Merkwirth et al., 2008
subunits of cytochrome c oxidase PHB1 function of the mitochondrial respiratory chain Tsutsumi et al., 2009
NADH-ubiquinone oxidoreductase 30 kDa subunit PHB1 stabilization of mitochondrial complex I Park et al., 2010
ND4 and ND5 PHB1 assembly of mitochondrial complex I Bourges et al., 2004
SLP-2 PHB1/2 stabilization of PHBs Da et al., 2008
TFAM PHB1 regulation of copy number of mitochondrial DNA Kasashima et al., 2008
ANT2 PHB2 unknown Kasashima et al., 2006
VDAC2 PHB2 unknown Kasashima et al., 2006
Hax-1 PHB2 stabilization of antiapoptotic Hax-1 Kasashima et al., 2006
nitric oxide-associated protein 1 (mNOA1) PHB1 regulation of mitochondrial protein translation and respiration Heidler et al., 2011
phosphorylated p66ShcA PHB1 putative protection against oxidative stress Madireddi, 2006

Transcription Factors

estrogen receptor PHB1/2 repression of estrogen-receptor activity Delage-Mourroux et al., 2000; He et al., 2008; Montano et al., 1999
E2Fs PHB1 repression of E2F activities Gamble et al., 2007; Joshi et al., 2003, 2007; Schneider et al., 2010
p53 PHB1 enhancement of p53-mediated transcriptional activity and chaperone activity of PHB1 for p53 Chander et al., 2011; Fusaro et al., 2003; Joshi et al., 2007
MyoD and MEF2 PHB2 repression of MyoD and MEF2 activities Sun et al., 2004, 2011
chicken ovalbumin upstream binding transcription factors I and II PHB2 transcriptional repression Kurtev et al., 2004
phosphorylated STAT3 PHB1 modulation of p53- and STAT3-mediated apoptosis Kathiria et al., 2012b

Other Nuclear Proteins

HDAC1 and HDAC5 PHB1/2 transcriptional repression Kurtev et al., 2004
Rb and its family members p107 and p130 PHB1 repression of E2F-mediated transcription and inhibition of cell proliferation Wang et al., 1999a, 1999b, 2002
Brg-1 and Brm PHB1 repression of E2F-mediated transcription and inhibition of cell proliferation Wang et al., 2002
heterochromatin protein 1 (HP1) family proteins PHB1 repression of E2F-mediated transcription and induction of cellular senescence Rastogi et al., 2006a
RING finger protein 2 (RNF2) PHB1/2 repression of the transcriptional activity of E2F1 and CP2c Choi et al., 2008; Lee et al., 2008
histone methyltransferase EZH2 PHB2 repression of estrogen-dependent transcription Hwang et al., 2008
PGC1α (PPARγ coactivator 1) PHB2 inhibition of the transcriptional activity of PPARγ Endo, 2011
minichromosomes maintenance complex of proteins MCM2, MCM5, and MCM 7 PHB1 inhibition of DNA replication Rizwani et al., 2009
nuclear receptor corepressor 1 (NCoR) PHB1 transcriptional repression of E2F family members Wang et al., 2002

Other Signaling Proteins

IgM receptor PBH1 phosphorylation of PHB1 Terashima et al., 1994
protease-activated receptor 1 PHB1/2 platelet aggregation Zhang et al., 2012
C-Raf PBH1 phosphorylation of C-Raf by Ras and consequently activation of the MEK-eIF4 pathway Chiu et al., 2012; Chiu et al., 2013; Rajalingam et al., 2005
Akt2 PHB2 cell-cycle exit and myogenic differentiation Héron-Milhavet et al., 2008; Sun et al., 2004
WD repeat domain 5 (WDR5, BIG3) PHB2 inhibition of the translocation of PHB2 to the nucleus Kim et al., 2009
Skp2B PHB1/2 attenuation of the transcriptional activity of p53, release of the repression of the estrogen receptor Chander et al., 2010, 2011; Umanskaya et al., 2007
MLK2 (mixed lineage kinase 2) PHB1 unknown Rasmussen et al., 1998
Shp1 (phosphatase) phospho-Y114-PHB1 modulation of insulin signaling Ande et al., 2009a
PKC-δ PHB1 phosphorylation of PHB1 leading to a decrease in the inner mitochondrial membrane permeability and to cell survival Zhu et al., 2010
14-3-3 protein PHB1 inhibition of PHB1 phosphorylation leading to an increase in the inner mitochondrial membrane permeability and to apoptosis Zhu et al., 2010
α-dystrobrevin PHB1 NB4 cell granulocytic differentiation Borutinskaite et al., 2011
transient Receptor Potential Melastatin 6 (TRPM6) PHB2 inhibition of TRPM6-mediated transepithelial Mg2+ transport Cao et al., 2009
HSP70 PHB1 resistance to stress Liu et al., 2010
glucose-regulated protein 75 (GRP75)/mortalin PHB1 cell proliferation, tumorigenesis and stress response in metastatic cells Martín et al., 2008
phospho- Ser291- Syk PHB1 B cell antigen-receptor signaling Paris et al., 2010
CRM-1 PHB1 export of PHB1 outside of the nucleus selectively in cancer cells Rastogi et al., 2006b
Pex14p PHB1 regulation of the peroxisomal importomer Oeljeklaus et al., 2012
annexin A2 PHB1/2 interaction blocked by calcium, association with lipid rafts Zhu et al., 2010
alpha-actinin PHB1/2 interaction blocked by calcium, association with lipid rafts Bacher et al., 2002
EHD2 Palmitoylated PHB1 inhibition of pyruvate carboxylase leading to a decrease in glucose and fatty acid oxidation Ande and Mishra, 2010; Vessal et al., 2006
C3 PHB1 enhancement of innate immunity response Mishra et al., 2007
CK1ε PHB2 inhibition of circadian transcription Kategaya et al., 2012

Proteins of Pathogens

SV40Tag (viral oncoprotein) PHB1 interruption of the association between PHB1 and Brg-1/Brm Wang et al., 2002
HIV-1 glycoprotein PHB1/2 viral spread in nonpermissive cells Emerson et al., 2010
DENV-2 E protein PHB2 internalization of the dengue virus Kuadkitkan et al., 2010
Chikungunya virus E2 protein PHB1 internalization of the chikungunya virus Wintachai et al., 2012
SARS-CoV nonstructural protein nsp2 PHB1/2 supposed disruption of intracellular host signaling during SARS-CoV infection Cornillez-Ty et al., 2009
Vi polysaccharide of Salmonella typhi PHB1/2 inhibition of the inflammatory response upon infection with Vi- S. typhi. Sharma and Qadri, 2004
capsid protein VP1 of foot-and-mouth disease virus PHB1 dephosphorylation of PHB T258 Chiu et al., 2012
macrophage surface HSP70 Leishmania donovani PHB1 induction of a strong humoral response in leishmaniasis patients Jain et al., 2010

See also Table S1.