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Flexibility in receptor usage by FMDV. Passage of FMDV in BHK-21 cells resulted in acquisition of amino acid replacements in the capsid, which expanded receptor usage (HS, heparan sulfate; integrin αvβ3; X, an unidentified receptor). Replacements are depicted as Van der Waals spheres in yellow (VP1, blue; VP2 green; VP3, red).
