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. 2020 Mar 9;375(1797):20190358. doi: 10.1098/rstb.2019.0358

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© 2020 The Authors.

Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.

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Figure 1. — Schematic of the conditions required to apply the Price equation. There are two sets of individuals, a and d, and some trait measurable on each individual. There are directed links from some individuals in set a to those in set d, which represent influence on the value of the trait. Fitness is defined, for individuals in set a, as the number of outgoing links. The trait values of the individuals in set d may or may not be equal to the average of the individuals they receive links from. (Online version in colour.)