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(a) minimal heparanase recognized sequence: R = H; SO3−; R1 and R2 uronic acids of Hep/HS chain. (b) Antithrombin binding site: R = SO3−; NAc R1 and R2 = uronic acids of Hep/HS chain; R3 = SO3; Fondaparinux: R = SO3−; R1 = H; R2 = Me; R3 = SO3−
