Table 1. The candidate genes chosen for functional analysis. *Crosses that were significantly different than their control counterpart using a standard Dunnett’s Test for statistical significance (P < 0.05). The genes are listed by p value from the GWAS.

Gene	GWAS p-value	Function
mas*	5.65E-06	serine protease, muscle attachment
CG32982	8.29E-06	Unknown function. Predicted role in actin cytoskeleton reorganization
Src64B*	1.01E-05	tyrosine-protein kinase,
bru1*	1.02E-05	RNA binding, alternative splicing, regulation of translation
Camta*	1.04E-05	DNA-binding transcription factor
dpr6	1.29E-05	Synapse organization, sensory perception
Hs6st	1.46E-05	Heparin sulfate 6-O-sulfotransferase, imaginal disc growth
CG32333	1.55E-05	Unknown function, predicted role in lipid metabolism
bab1	2.30E-05	DNA-binding transcription factor, pigmentation, sex differentiation
tey*	2.32E-05	E3 ubiquitin-protein ligase, negative regulation of transcription
mtgo*	2.32E-05	Unknown function, essential.
Asph	2.78E-05	Aspartyl β-hydroxylase that may have a role in neurogenesis
Cpr66D	3.0E-05	Structural constituent of cuticle
CG15096	3.78E-05	Transmembrane transport
app	4.49E-05	Palmitoyltransferase involved in planar cell polarity
zormin*	5.52E-05	Protein found in the Z-disc and the M-line of muscles. Likely structural function
fru	6.82E-05	zinc finger transcription factor important for sexual differentiation
Lmpt	7.28E-05	Zinc ion binding, defense response
bru3*	7.84E-05	RNA binding, negative regulation of translation, regulation of alternative mRNA splicing