TABLE 2.

Test results for homogeneity of multivariate dispersions (PERMDISP) and pairwise permutation tests using the betadisper and permutest functions in vegan.

Test	Comparison	Test statistic	p-value
PERMDISP	Site	0.5837	ns
	Depth	11.565	0.0001
Permutation test	10 m vs. 16 m	–0.1384	ns
	10 m vs. 25 m	3.9797	0.0004
	10 m vs. 35 m	4.2363	0.0001
	16 m vs. 25 m	4.0923	0.0001
	16 m vs. 35 m	4.3452	0.0001
	25 m vs. 35 m	0.3414	ns

Pairwise p-values are the permuted p-values generated by the test (9,999 permutations). Non-significant tests are listed as “ns.”