Table 1.

Formulas applied to calculate networks parameters

Measure	Formula	Reference
Degree centrality (D)	$D=\sum _{i=1}^{n}a(p_i,p_k)$	Freeman (1978)
Betweenness centrality (B)	$B=\sum _i^n\sum _{<j}^n\frac{g_{\mathit{ij}}(p_k)}{g_{\mathit{ij}}}$	Freeman (1978)
Closeness centrality (C)	$C=\frac{n-1}{{\sum }_{i=1}^n{n}_d(p_i,p_k)}$	Freeman (1978)
Density (ρ)	$\rho \frac{2E}{n(n-1)}$	Martínez‐López et al. (2009)
Diameter (Ø)	maxik d(i, k)	West Douglas (2001)

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