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. 2020 Apr 22;929:117–161. doi: 10.3897/zookeys.929.50240

Table 1.

Species groups of Hipposideros (modified from Murray et al. 2012 to include newly recognized forms and to remove species now recognized in Doryrhina and Macronycteris).

Armiger group calcaratus subgroup H. macrobullatus H. lankadiva
H. alongensis H. calcaratus c H. maggietaylorae H. lekaguli
H. armiger H. cervinus c H. nequam H. pelingensis
H. griffini a H. coxi c H. obscurus Larvatus group
H. pendelburyi a H. galeritus c H. orbiculus H. grandis
H. turpis ruber subgroup H. papua H. khasiana” a,g
Bicolor group H. abae d H. pygmaeus H. larvatus
ater subgroup H. beatus e Boeadii group H. madurae
H. ater b H. caffer e H. boeadii H. sorenseni
H. atrox a H. fuliginosus e Cyclops group f H. sumbae
H. bicolor b H. lamottei e H. corynophyllus Megalotis group
H. breviceps b H. ruber e H. edwardshilli H. megalotis
H. cineraceus b H. tephrus a,e H. muscinus Pratti group
H. coronatus b subgroup uncertain H. semoni H. lylei
H. dyacorum b H. cruminiferus H. stenotis H. pratti
H. einnaythu a,b H. curtus H. wollastoni H. scutinares
H. halophyllus b H. doriae Diadema group Speoris group
H. khaokhouayensis b H. durgadasi H. demissus H. jonesi h
H. nicobarulae a,b H. fulvus H. diadema H. marisae g
H. pomona b H. gentilis a H. dinops H. speoris
H. ridleyi b H. hypophyllus H. inexpectatus
H. rotalis b H. kunzi a H. inornatus

(Endnotes) a Added to species list subsequent to Murray et al. (2012) b Recognized in the Ater species group by Monadjem (2019) c Recognized in the Calcaratus species group by Monadjem (2019) d Formerly listed in the Speoris group but transferred to the Ruber group by Monadjem (2019) e Recognized in the Ruber species group by Monadjem (2019) f H. cyclops and H. camerunensis are now recognized as members of Doryrhina; listed species were treated as Doryrhina inMonadjem (2019) on the basis of similar morphology but were recognized as the Muscinus group by Tate (1941); they might represent an unnamed genus or subgenus. g Invalid name accorded to what is likely a real biological entity (cf. Monadjem 2019) h Formerly in the Bicolor species group but transferred to the Speoris group by Monadjem (2019).