Table 1.
Armiger group | calcaratus subgroup | H. macrobullatus | H. lankadiva |
H. alongensis | H. calcaratus c | H. maggietaylorae | H. lekaguli |
H. armiger | H. cervinus c | H. nequam | H. pelingensis |
H. griffini a | H. coxi c | H. obscurus | Larvatus group |
H. pendelburyi a | H. galeritus c | H. orbiculus | H. grandis |
H. turpis | ruber subgroup | H. papua | “H. khasiana” a,g |
Bicolor group | H. abae d | H. pygmaeus | H. larvatus |
ater subgroup | H. beatus e | Boeadii group | H. madurae |
H. ater b | H. caffer e | H. boeadii | H. sorenseni |
H. atrox a | H. fuliginosus e | Cyclops group f | H. sumbae |
H. bicolor b | H. lamottei e | H. corynophyllus | Megalotis group |
H. breviceps b | H. ruber e | H. edwardshilli | H. megalotis |
H. cineraceus b | H. tephrus a,e | H. muscinus | Pratti group |
H. coronatus b | subgroup uncertain | H. semoni | H. lylei |
H. dyacorum b | H. cruminiferus | H. stenotis | H. pratti |
H. einnaythu a,b | H. curtus | H. wollastoni | H. scutinares |
H. halophyllus b | H. doriae | Diadema group | Speoris group |
H. khaokhouayensis b | H. durgadasi | H. demissus | H. jonesi h |
H. nicobarulae a,b | H. fulvus | H. diadema | H. marisae g |
H. pomona b | H. gentilis a | H. dinops | H. speoris |
H. ridleyi b | H. hypophyllus | H. inexpectatus | |
H. rotalis b | H. kunzi a | H. inornatus |
(Endnotes) a Added to species list subsequent to Murray et al. (2012) b Recognized in the Ater species group by Monadjem (2019) c Recognized in the Calcaratus species group by Monadjem (2019) d Formerly listed in the Speoris group but transferred to the Ruber group by Monadjem (2019) e Recognized in the Ruber species group by Monadjem (2019) f H. cyclops and H. camerunensis are now recognized as members of Doryrhina; listed species were treated as Doryrhina inMonadjem (2019) on the basis of similar morphology but were recognized as the Muscinus group by Tate (1941); they might represent an unnamed genus or subgenus. g Invalid name accorded to what is likely a real biological entity (cf. Monadjem 2019) h Formerly in the Bicolor species group but transferred to the Speoris group by Monadjem (2019).