(a) Cortical ripples in the retrosplenial cortex (RSC). Exemplary trace showing a ripple recorded in the CA1 and eight simultaneously recorded channels in the retrosplenial cortex ranging from deep (top, light grey) to superficial (bottom, black). Note that retrosplenial ripple is localized to superficial layers. Bottom channel is overlaid with a low-passed filtered version (dashed line) to illustrate the cortical sharp wave, in which the ripple is embedded. (b) Ripple oscillations in the MTL are coupled to cortical ripples in temporal association area, but not in the primary cortex. HPC, hippocampal cortex. From Vaz et al. [117], with permission. (Online version in colour.)