An integrative, three-pronged study of olfactory communication about reproductive state in the Coquerel's sifaka (Propithecus coquereli). Shown are differences in (a) chemical design, based on principal component and linear discriminant analyses of the chemical compounds identified in female genital secretions, using gas chromatography and mass spectrometry; (b) the frequency of signal delivery in a forest enclosure, based on observed urogenital marking by a representative female; and (c) the perception of or interest in signals across reproductive seasons, based on conspecific investigation, in situ, of freshly deposited marks. Adapted from Greene & Drea [84]. *p < 0.05; ***p < 0.001.