BLH2 and BLH4 modulate the expression of PME-related genes in seeds. A and B, Expression analysis of PME genes (A) and genes encoding modulators of PME (B) in wild-type (WT) and blh2 blh4 seeds at 7 to 10 DPA. C, Expression of CESA5, MUM5, SOS5, and FEI2 in wild-type and blh2 blh4 seeds at 7 to 10 DPA. Data represent means ± sd of three biological replicates. Asterisks denote significant differences from the wild type based on one-way ANOVA analysis (**P < 0.01).