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. 2020 May 8;10:7785. doi: 10.1038/s41598-020-64563-7

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Grooming behavior of Halobates. (A–D) Different stages of grooming exhibited by Halobates hayanus with specialized activities. (A) Tibio-tarsal regions of the fore legs are held together and rubbed against each other; (B) One fore leg brushes over the posterior region of the middle leg; (C) Posterior regions of middle and hind legs brush against each other with the body supported by the other legs; (D) Fore legs brush the ventral abdominal metathoracic region alternately. (E–H) Transfer of waxy secretion (shown in blue) from metathoracic gland to various parts of the body.