TABLE 3.

Phylogenetic relatedness of bacteria and archaea grouped by life-history strategy.

Location	Depth	Life history strategy	#OTUs	NRI	NTI
Hillslope	0 to 5 cm	Winter-adapted	55	−0.48	2.18
		Snowmelt-specialists	335	−2.15	1.13
		Spring-adapted	1093	5.28	2.75
	5–15 cm	Winter-adapted	138	−1.67	5.16
		Snowmelt-specialists	433	−0.89	0.33
		Spring-adapted	1120	9.53	5.21
	15 cm +	Winter-adapted	132	1.69	3.37
		Snowmelt-specialists	461	−0.16	0.31
		Spring-adapted	771	1.77	0.47
Floodplain	0 to 5 cm	Winter-adapted	83	2.05	3.49
		Snowmelt-specialists	269	3.13	2.25
		Spring-adapted	631	0.05	0.19
	5–15 cm	Winter-adapted	123	−0.24	3.32
		Snowmelt-specialists	153	3.48	3.88
		Spring-adapted	387	3.63	−0.01
	15 cm +	Winter-adapted	119	1.60	−1.27
		Snowmelt-specialists	200	3.49	3.07
		Spring-adapted	276	1.32	1.00

Values in bold indicate a significant pattern of phylogenetic relatedness. Positive NTI/NRI values indicate that groups are phylogenetically clustered, with a low probability that the observed phylogenetic distance within groups is greater compared to the null model (P < 0.025). Negative NTI/NRI values indicate phylogenetic overdispersion, with a high probability that the observed phylogenetic distance within groups is greater than the null model (P > 0.975).