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. 2020 May 25;13:4649–4663. doi: 10.2147/OTT.S252990

Table 7.

Systematic Review of Genetic Alterations at the PTEN and CDKN2A Loci in Chordomas

No. Author (Year) Materials/Objects Method Findings
1 Mertens et al (1994)11 Human sacral chordoma samples (n=3) Chromosome banding analysis Karyotype included del(9) and –10 in one case.
2 Scheil et al (2001)4 Human chordoma samples (n=16); U-CH1 cell line Comparative genomic hybridization; chromosome banding analysis; FISH Loss of 9p was identified in 5 of 16 (31%) cases, and loss of chromosome 10 segment was noted in 3 of 16 (19%) cases; U-CH1 had del(9) and del(10) chromosomal abnormalities.
3 Kuźniacka et al (2004)5 Human chordoma samples (n=7) Chromosome banding analysis Loss of chromosome 10 segment was noted in 3 of 7 (43%) cases, and loss of 9p sequences was identified in 2 of 7 (29%) cases.
4 Hallor et al (2008) Human chordoma samples (n=26) Array comparative genomic hybridization; FISH Heterozygous deletion at 9p21 covering the CDKN2A locus was observed in 15 of 26 (58%) tumor samples, homozygous deletion was found in 3 of 26 (12%) samples.
5 Han et al (2009)34 Human chordoma samples (n=11), and U-CH1 cell line Immunohistochemistry; western-blotting Weak PTEN staining was demonstrated in 4 of 11 (36%) cases, and 6 of 11 (55%) cases were negative for PTEN staining, whereas 1 (9%) case showed positive PTEN staining; PTEN expression was not observed in U-CH1 cells.
6 Presneau et al (2009)35 Human chordoma samples (n=50) Immunohistochemistry PTEN displayed no immunoreactivity in 7 of 43 (16%) chordoma samples, and CDKN2A showed no immunoreactivity in 46 of 48 (96%) chordoma samples.
7 Sommer et al (2010)36 Human chordoma samples (n=27) Immunohistochemistry CDKN2A expression was absent in 20 of 27 (74%) chordomas. There was no significant difference in median disease-free survival between CDKN2A-positive chordomas (90 months) and negative chordomas (24 months).
8 Horbinski et al (2010)13 Human clival chordoma samples (n=28) FISH; PCR-based microsatellite LOH analysis 9p LOH was identified in 3 of 25 (12%) samples, and 10q LOH was found in 12 of 21 (57%) samples; FISH identified homozygous loss of the 9p21 locus in 5 of 23 (22%) cases.
9 Brüderlein et al (2010)7 U-CH1 and U-CH2 chordoma cell lines Array comparative genomic hybridization Both U-CH1 and U-CH2 showed biallelic loss of the CDKN2A locus on chromosome 9p21; loss of the PTEN locus was noted in U-CH1, but not in U-CH2.
10 Shalaby et al (2010)37 Human chordoma samples (n=147) Immunohistochemistry PTEN expression was absent in 19 of 147 (13%) analyzed chordomas.
11 Le et al (2011)6 Human chordoma samples (n=20) Array comparative genomic hybridization Loss of CDKN2A was observed in 16/20 (80%) cases, six of which showed homozygous deletion; loss of PTEN was found in 16/20 (80%) cases; no mutations were found in any of the 21 chordoma samples.
12 Dewaele et al (2011)14 Human chordoma samples (n=42) FISH Loss of PTEN was observed in 7 of 18 (39%) analyzed samples.
13 Rinner et al (2012)9 MUG-Chor1 chordoma cell line Affymetrix GeneChip Human Mapping SNP 6.0 array The cell line displayed chromosomal loss at 10p15.3-q23.32 (includes the PTEN locus), and a homozygous loss was seen at 9p24.3-p13.1 (includes the CDKN2A locus).
14 Diaz et al (2012)17 Human chordoma samples (n=22) Genome-wide high-resolution SNP-array Loss of 9p involving CDKN2A was identified in 4 of 18 (22%) chordoma samples.
15 Choy et al (2014)8 Human chordoma samples (n=45) Matrix-assisted laser desorption/ionization time-of-flight mass spectrometer A nonsense mutation (p.R130*) in PTEN was identified in 1 of 45 (2.2%) chordoma samples; a nonsense mutation (p.R58*) was identified in CDKN2A in 2 of 45 (4.4%) chordoma samples.
16 Chen et al (2014)38  Human sacral chordoma samples (n=40) Immunohistochemistry Positive expression of PTEN was observed in 10/40 (25%) chordoma samples; patients with negative PTEN expression had a poorer prognosis than those with positive PTEN expression (P=0.011).
17 Lee et al (2015)16 Human chordoma samples (n=23) PCR-based microsatellite LOH analysis LOH was observed at 10q23 (includes the PTEN locus) in 6 of 23 (26%) specimens; chordoma tissues with LOH at 10q23 showed decreased or absent PTEN expression, and those without LOH at 10q23 demonstrated positive PTEN expression.
18 Wang et al (2016)18 Human chordoma samples (n=24) Genome-wide high-resolution SNP-array Copy number loss of CDKN2A was identified in 12 of 22 (55%) chordoma samples, and homozygous deletion of CDKN2A was detected in 5 of 22 (23%) samples.
19 Cottone et al (2020)15 Human chordoma samples (n=384) Immunohistochemistry; FISH Complete loss of chromosome 9 was observed in 3 cases; Positive CDKN2A protein expression was noted in 141 of 303 (47%) samples; copy number loss of CDKN2A gene was found in 138 of 274 (50%) samples, including heterozygous loss in 44 samples and homozygous loss in 48 samples.

Abbreviations: FISH, fluorescent in situ hybridization; LOH, loss of heterozygosity.