TABLE 1.

Abiotic and biotic factors at different scales.

Factors	Scales	Explanations
Abiotic
Site	Site (large-scale)	Effects of this treatment are related to abiotic factor changes at regional scale, here, the effects are mainly due to the variance of soil pH and abiotic hydrothermal conditions.
Habitat Habitat × site	Microhabitat (small-scale)	Effects of these treatments are related to abiotic changes in meso-topography, here, the effects are mainly due to the variance of soil nutrients.
Biotic
Cushion Cushion × site	Cushion presence	Effects of these treatments are relevant to cushion presence/absence (i.e., differences between cushion and open microhabitats).
Cushion × habitat cushion × habitat × site	Cushion phenotype	Effects of these treatments are relevant to cushion phenotypes (i.e., loose- and tight-cushions).