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. 2020 May 28;11:567. doi: 10.3389/fpls.2020.00567

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Copyright © 2020 Renzaglia, Browning and Merced.

This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

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Leucobryum glaucum. LM longitudinal sections of astomate capsule. (A) Base of immature capsule where seta meets the neck covered by calyptra (C). An inconspicuous fluid-filled intercellular space (IS) extends the entire length of the region between the amphithecium that forms the capsule wall, and the endothecium (En) that consists of a prominent columella (Co) and developing spore sac with one layer of archesporium (A) (sporogeneous tissue). (B) Fully expanded capsule. With development of the spore sac that contains 100s of spores (S), the columella (Co) has partially degenerated and the intercellular spaces are closed (arrow) or residual (IS). Bars = 25 μm.