Evidence of horizontal symbiosis gene transfer between genospecies. (a)–(c) Examples of symbiosis-gene phylogenies, with insets showing clades in which identical alleles are shared across genospecies. (d) Distribution of plasmid groups, which were defined based on the genetic similarity of the RepA plasmid partitioning protein. (e) Phylogenetic analysis of the repA gene of plasmid type Rh08. A complete set of conjugal transfer genes has the following genes upstream of repA: traI,trbBCDEJKLFGHI,traRMHBFACDG, with the origin of transfer (oriT) between traA and traC. Partial sets are broken by the end of the scaffold, mostly after traM. The colour of the diamond indicates the genospecies origin with reference to (d).