Morphospecies of Chasmosaurus (A–D) and Pentaceratops (E–J) clades which do not overlap stratigraphically. These are hypothesized to form two anagenetic lineages which resulted from a cladogenetic branching event prior to the middle Campanian. (A) “Chasmosaurus russelli”, lower Dinosaur Park Fm, ~76.8 Ma. (B) Chasmosaurus belli, middle Dinosaur Park Fm, ~76.5–76.3 Ma. (C) Vagaceratops irvinensis, upper Dinosaur Park Fm, ~76.1 Ma. (D) Kosmoceratops richardsoni, middle Kaiparowits Fm, ~76.0–75.9 Ma. (E) Utahceratops gettyi, middle Kaiparowits Fm, ~76.0–75.6 Ma. (F), c.f. Pentaceratops sternbergii, unknown occurrence within “Fruitland Formation” ~76.0–75.1 Ma. (G) Aff. Pentaceratops n. sp., uppermost Fossil Forest Mbr, Fruitland Fm, ~75.1 Ma. (H) Navajoceratops sullivani, lowermost Hunter Wash Mbr, Kirtland Fm, ~75.0 Ma. (I) Terminocavus sealeyi, middle Hunter Wash Mbr, Kirtland Fm, ~74.7 Ma. (F) Anchiceratops ornatus, Drumheller to Morrin Mbr, Horseshoe Canyon Fm, ~71.7–70.7 Ma. Stratigraphic positions and recalibrated radiometric dates from Supporting Information 1 and Fowler (Chapter 2). Timescale from Gradstein et al. (2012). Specimens not to scale. Images adapted from Lehman (1998); Holmes et al., 2001; Sampson et al. (2010); Maidment & Barrett (2011); and Longrich (2014).