Phenotypic variation in Papilio dardanus and samples used. (a) Seven female forms and a male. (b) Origin of samples for sequencing and population genetic analyses, from four subspecies: P. dardanus polytrophus (Kenya), P. dardanus tibullus (Kenya), P. dardanus cenea (South Africa), and P. dardanus meriones (Madagascar). The specimen of subspecies P. dardanus tibullus was used for the construction of the draft genome sequence. The tree depicts the relationships among these four subspecies and is based on a tree presented in [3]. Three female forms were analysed: hippocoonides, cenea, and ‘male-like’.