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. 2020 Jun 9;10:9284. doi: 10.1038/s41598-020-65999-7

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Manhattan plots for GWAS of the first principal component for the hyperspectral region 707–922 nm for environments TX08, AR09 and AR10. Only non-pigmented bran accessions are presented in this analysis. The x-axis displays chromosome pseudomolecule coordinates and the y-axis displays the –log₁₀(p) value for each SNP across chromosomes. The dark horizontal line represents the genome-wide significance threshold. The labeled SNPs are discussed as potential candidates in the text. GW5 – grain weight 5 gene; LTPL164 – Protease inhibitor/seed storage gene.