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. 2020 Jun 10;10:9393. doi: 10.1038/s41598-020-66229-w

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Comparison of cancer stem cell markers, SOX2 and CD44 between BC 2.5D organoid at the early and late passage and their original BC 3D organoids. Dog urothelial carcinoma cells were used as 2D cell lines. Expression level of SOX2 and CD44 in 3D and 2.5D organoids from different strains (BC1; A and BC2; B) and 2D cell lines was analyzed by quantitative real-time PCR (n = 4) and quantified based on the ratio of expression level to GAPDH. Data were expressed as mean ± SEM. *P < 0.05 vs. 3D organoid.