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. Author manuscript; available in PMC: 2021 Aug 1.
Published in final edited form as: Sleep Med Rev. 2020 Mar 17;52:101307. doi: 10.1016/j.smrv.2020.101307

TABLE 2:

Summary of 10 studies on CSF hypocretin-1 and metabolic function in children, women and men

Author, Year Country Research Question Study Design Study population Exposure assessment Outcome (s) assessment HCRT1 Assessment Main Findings
Arnulf, 2006, USA [54] Association between HCRT 1 and leptin Cross-sectional 93 narcoleptic low HCRT1
72 narcoleptic normal HCRT1
89 w/other sleep disorders
111 controls
HCRT1 deficiency Serum, CSF leptin, CRP Cerebrospinal fluid
Radio-immunoassay
Serum leptin similar in all groups;
HCRT 1 was not associated with serum or CSF leptin;
CRP levels similar across groups;
Analyses adjusted for sex, age, BMI, CRP, sleep duration, meds use
Beitinger 2011, Germany [70] Association between HCRT1 deficiency and glucose metabolism Case-control 17 adults with NC
17 controls
HCRT1 deficiency Glucose, insulin sensitivity following 4-hour oral glucose tolerance test Cerebrospinal fluid
Radio-immunoassay
No differences in glucose metabolism between groups;
Study matched for age, sex and BMI
Donadio 2014, Italy [74] Association between HCRT 1, sympathetic and cardiovascular function Cross-sectional 19 adults with NC
19 controls
HCRT1 deficiency HR, BP Cerebrospinal fluid
Radio-immunoassay
HCRT1 deficiency is associated with lower HR and BP;
HCRT 1 correlated (positively) with HR, but not with BP;
Study matched for age, sex
Fronczek 2008, Holland [72] Association between HCRT1 and autonomic balance Cross-sectional 15 men with NC
15 controls
HCRT1 deficiency Resting MET rate, HR, BP Cerebrospinal fluid
Radio-immunoassay
Similar resting MET rate, mean HR, systolic and diastolic BP among NC men and controls, but variability of HR and BP differed between groups;
Study matched for sex, BMI, age
Heier 2011, Norway [67] Association between HCRT 1 and metabolic risk factors Cross-sectional 26 adults with NC, deficient HCRT1
23 adults with NC, normal HCRT1
11 adults with IH
43 controls
HCRT1 deficiency BMI, HbA1C, serum and CSF leptin Cerebrospinal fluid
Radio-immunoassay
Similar mean BMI in all groups;
serum, CSF leptin not associated with HCRT 1 levels;
similar HbA1C across all groups;
Study matched for age, sex
Omokawa 2016, Japan [68] Associations between HCRT1 and BMI Cross-sectional 14 patients with PWS
37 with NC
14 with IH
HCRT1 BMI Cerebrospinal fluid Radio-immunoassay Moderate HCRT1 in PWs significantly higher than NC patients, but lower than those with IH. HCRT 1 and BMI not correlated;
Study controlled for race/ethnicity
Poli 2009, Italy [65] Associations between HCRT 1, metabolic function and MET syndrome Cross-sectional 14 men with NC
14 men with IH
HCRT1 deficiency Adiposity measures, systolic, diastolic BP, HR, lipid profile, glucose, insulin Cerebrospinal fluid
Radio-immunoassay
NC men had higher adiposity, BP, total cholesterol, triglycerides, insulin, leptin, but similar HR, lower HDL vs. IH men;
MET syndrome only present in NC group (64%);
Study controlled for age and sex;
all patients were white and drug naive
Poli 2013, Italy [66] Associations between HCRT 1deficiency, precocious puberty and BMI Cross-sectional 43 children with NC
52 obese controls
HCRT1 deficiency Adiposity measures, lipid profile, CRP, glucose, insulin, precocious puberty Cerebrospinal fluid
Radio-immunoassay
HCRT1 lower in NC group with vs. without precocious puberty and with excessive vs. normal weight, statistically insignificant;
Precocious puberty shown in 17% and 2% of NC patients and controls, respectively;
Study matched for age
Santiago 2018, Germany [69] Association between HCRT1 and body composition Cross-sectional 26 adults BMI≥25
15 adults normal weight
HCRT1 BMI, body weight, fat, fat-free mass, total water, WC Cerebrospinal fluid Radio-immunoassay Inverse associations between HCRT 1, body weight, WC, fat-free mass and total water;
no association with BMI, fat mass, insulin, glucose and adiponectin;
Study matched for race/ethnicity
Sieminski 2017, Finland [73] Associations between HCRT1 deficiency and BP patterns Cross-sectional 11 adults with NC, HCRT1 depleted
10 adults with NC, HCRT1 deficient
HCRT1 deficiency BP patterns, daytime and nocturnal systolic, diastolic BP, Dip of systolic, diastolic BP Cerebrospinal fluid
Radio-immunoassay
Nocturnal, daytime BP patterns similar in both groups;
No information about control for confounding

BMI=body mass index; BP=blood pressure; CSF=cerebrospinal fluid; HCRT1=hypocretin-1; HR=heart rate; IH=idiopathic hypersomnia; MS=metabolic syndrome; NC=narcolepsy and cataplexy; PWS=Prader-Willi Syndrome; WC=waist circumference;