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. 2020 Jul 22;10:12171. doi: 10.1038/s41598-020-68739-z

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Metabolites negatively correlated with freezing behavior are also observed in human AD urine samples. (a–c) Graphs representing the top metabolites positively and negatively correlated with freezing behavior across ages in the PFC, HPC, and spleen of Ctrl mice. (d–f) Graphs representing the top metabolites positively and negatively correlated with freezing behavior across ages in the PFC, HPC, and spleen of AD mice. The red bars denote negative correlations and the green bars denote positive correlations to freezing behavior. The red stars indicate metabolites that are changed in AD human urine samples when compared to a library using MetaboAnalyst. Ctrl control, AD Alzheimer’s disease, PFC prefrontal cortex, HPC hippocampus.