Nonlinear and interaction responses to environment. (A) Nonlinear response of species to a temperature gradient () is fitted with a model containing a quadratic term, . (B) Combined effects of temperature and land cover () are fitted with an additional interaction term, , where response to temperature might differ in forests and fields. (C) Alternatively, nonlinear and interaction responses enter indirectly though products with other species (blue and brown) that depend on temperature , but require the time dimension. The background shading in C describes Hutchinson’s concept of the fundamental niche that might be realized by species in the absence of other species.