Table 1.
KSHV modulation of the host immune system.
Responses | ORF | Gene product | Celluar homolog | Functions | Mechanisms |
---|---|---|---|---|---|
Complement cascade | ORF4 | KCP | RCA | Blocks complement activation | Enhances C3 convertase decay; prevents C3b deposition; factor I cofactor activity |
IFN signaling | K9 | vIRF1 | IRF | Inhibits IRF3-mediated transcription; cell transformation | P300/CBP sequestration; antagonizes p53, ATM, GRIM 19 activity and TGFβ signaling? |
K11.1-K11 | vIRF2 | IRF | Inhibits IRF3-mediated transcription | ||
K10.5-K10.6 | vIRF3 | IRF | Inhibits IRF7-mediated transcription | Inhibition of IRF7 DNA binding | |
ORF50 | RTA | Inhibits IRF7 activation | IRF7 ubiquitination and degradation | ||
ORF45 | Inhibits IRF7 activation | Inhibition of IRF7 phosphorylation/nucleartranslocation | |||
K8 | K-bZIP | Inhibits IRF3-mediated transcription | Transcription repressor of IFN-β promoter | ||
Chemokine Network | K4 | vCCL2 | CC chemokine | Th2 chemotaxis, angiogenesis | CCR3 agonist; C-, CC-, CXC-, CX3C-R antagonist |
K6 | vCCL1 | CC chemokine | Th2 chemotaxis, angiogenesis | CCR8 agonist | |
K4.1 | vCCL3 | CC chemokine | Th2 chemotaxis, angiogenesis | CCR4 agonist | |
ORF74 | vGPCR | GPCR | Inflammation, angiogenesis; cell transformation | Activates PI3K/AKT/mT0R pathway; activates NFkB, AP-1, HIF-1α trascriptional activities | |
K2 | vIL6 | IL6 | Inflammation, angiogenesis | IFNa-inducible; gp80-independent signaling; activates JAK/STAT and MAPK pathway | |
K12 | Kaposin B | Increases cytokines release | Activates MK2, blocks cytokine mRNA decay | ||
K14 | vCD200 | CD200 | Suppress macrophage activity | CD200R-mediated inhibitory signaling | |
Apoptosis and autophagy | K13 | vFLIP | FLIP | Anti-apoptosis, in cell transformation spindle shape, inflammation | Blocks activation of caspase 8? NFκB pathway activation |
ORF16 | vBcl-2 | Bcl-2 | Anti-apoptosis, anti-autophagy | Inhibits pro-apoptotic Bcl-2 family proteins; inhibits Beclin1-mediated autophagy | |
K7 | vIAP | IAP | Anti-apoptosis, interferes with ubiquitin-proteasome pathway | Inhibits caspase 3 and CAML activity; inhibits PLIC-regulated Ub-proteasome pathway | |
ORF73 | LANA | Cell growth transformation | Inhibits p53, Rb, GSK-3β; and TGF-β pathways | ||
ORF50 | RTA | Anti-apoptosis | Inhibits p53 transcriptional activity | ||
CTL and NK responses | K3 | vMIR1 | MARCH | CTL cell evasion | Downregulation of HLA-A, -B, -C, and -E |
K5 | VMIR2 | MARCH | CTL and NK cell evasion | Downregulation of HLA-A, -B, ICAM-1, B7–2, MICA/B, ALCL, CDld, PECAM, ALCAM and IFNγRI |