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. 2020 Jul 28;15(7):e0226934. doi: 10.1371/journal.pone.0226934

Table 1. Descriptions of hornet aggression behaviors observed and their corresponding scores.

Behavior Description Intensity score
Win Loss Tie Win Lose Tie
Threatening Rushing opponent Falling back or escaping Confrontation and separation 2 0 1
Lifting antennae and front legs and shaking wings, (Fig 1B) Falling back or escaping Confrontation 2 0 1
Opening mandible, (Fig 1F) Falling back or escaping Confrontation 2 0 1
Threateningly flying over opponent Escaping Disregard or confrontation 2 0 1
Chasing opponent Escaping Confrontation and separation 2 0 1
Grappling Pushing or fighting while flying Falling back or escaping Confrontation and separation 3 0 1
Banging or pushing with head Falling back or escaping Confrontation and separation 3 0 1
Chasing and grabbing, (Fig 1E) Being thrown off Confrontation and separation 3 0 1
Forcing down and throwing opponent, (Fig 1D) Being thrown off Confrontation and separation 3 0 1
Getting opponent and biting or stinging, (Fig 1A) Being bitten or stung Confrontation and separation 4 0 1
Killing Killing opponent, (Fig 1C) Being killed Failure to kill opponent or both sides dying 5 0 1