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. 2020 Jul 6;10(14):7686–7712. doi: 10.1002/ece3.6492

TABLE 1.

List of the 27 variables used in the current study. Each landcover class‐level metric has been calculated for each land type. Abbreviations for categories: Bare Land (Br), Grassland (Gr), Needle‐leaved forest (NLF), Shrubland (Shr), Snow and Ice (Sn)

Category Variable Abbreviation Layer Source
Linear and point features Density of highways and national roads DENS_rd Berman, 2009
Density of human settlements DENS_set OpenStreetMap
Density of rivers DENS_riv www.DIVA‐GIS.org
Topographic Slope position SLP
Focal mean of elevation ELEV
Roughness ROUGH NASA’s SRTM v.4 (Jarvis et al., 2008)
Compound topographic index CTI
Dissection DISS
Climatic Annual mean temperature TEMP WorldClim Version 2 (Fick & Hijmans, 2017)
Landcover (Landscape level) Aggregation index AI
Contrast‐weighted edge index CWED ESA GlobCover 2009 v2.3 (Arino et al., 2008)
Patch density PD
Landcover (Class Level) Area‐weighted mean AREA_AM_(Br, Sn, Gr, Shr, NLF)
Percentage of landscape PLAND_(Br, Sn, Gr, Shr, NLF) ESA GlobCover 2009 v2.3 (Arino et al., 2008)
Radius of gyration (mean area) GYR_AM_(Br, Sn, Gr, Shr, NLF)