Maximum-likelihood phylogenetic tree of DENV-1 detected in a traveler who returned from Benin to France (red circle) along with other strains from Africa (bold), strain provided by the French National Reference Centre for Arbovirus (green square), and reference strains. The general time-reversible model (discrete γ distribution with evolutionarily Invariant sites) was used to construct the tree with 130 full-genome sequences. Bootstrap support values (percentage of 500 replicates) are shown at nodes. The tree was rooted with reference strains of DENV-2, DENV-3, and DENV-4 (not shown). Scale bar indicates nucleotide substitutions per site. DENV, dengue virus.