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. 2020 Jul 29;13:133. doi: 10.1186/s13068-020-01773-1

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© The Author(s) 2020

Open AccessThis article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated in a credit line to the data.

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Fig. 6 — Effect of lipid modifications on the efflux of lupeol during two-phase fermentation. a Schematic representation of two-phase extractive fermentation. b Ratio of the extra/intracellular lupeol content after 120 h of fermentation, calculated based on the lupeol titer as shown in Additional file 1: Fig. S5. c Effect of lipid modifications on cell growth. d Lupeol productivity after 120 h of fermentation. These results indicated that the efflux of lupeol was enhanced and ultimately improved cell growth and the capacity of lupeol synthesis in engineered strains. Error bars represent ± SD of biological triplicates