Table 2.
Characteristic |
1 |
2 |
3 |
4 |
5 |
6 |
7 |
8 |
9 |
10 |
11 |
12 |
13 |
14 |
15 |
16 |
17 |
18 |
19 |
20 |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
Motility |
+ |
– |
+ |
+ |
+ |
+ |
– |
+ |
+ |
+ |
+ |
+ |
+ |
– |
+ |
+ |
– |
+ |
+ |
|
Oxidase |
+ |
+ |
+ |
+ |
+ |
+ |
V |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
– |
+ |
+ |
+ |
+ |
+ |
Catalase |
+ |
+ |
+ |
w |
V |
+ |
– |
+ |
+ |
– |
+ |
+ |
M |
+ |
V |
– |
+ |
– |
+ |
+ |
Urease |
– |
+ |
+ |
– |
V |
– |
– |
+ |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
Nitrate reduction |
+ |
– |
+ |
+ |
V |
+ |
F |
M |
+ |
+ |
+ |
+ |
M |
+ |
M |
+ |
V |
– |
+ |
+ |
Hippurate hydrolysis |
– |
– |
– |
+ |
– |
– |
– |
– |
– |
F |
– |
– |
– |
+ |
– |
– |
M |
– |
– |
– |
Indoxyl acetate hydrolysis |
+ |
– |
+ |
+ |
– |
+ |
– |
V |
+ |
V |
– |
– |
– |
– |
M |
+ |
+ |
– |
– |
– |
Alkaline phosphatase |
– |
– |
+ |
– |
– |
– |
M |
+ |
– |
V |
– |
– |
– |
– |
– |
– |
U |
– |
– |
F |
H2S production TSI |
– |
U |
+ |
– |
V |
– |
– |
+ |
– |
F |
– |
– |
– |
– |
– |
– |
– |
– |
+ |
+ |
Alpha haemolysis |
+ |
– |
– |
– |
– |
F |
F |
– |
+ |
F |
– |
– |
V |
– |
– |
+ |
– |
– |
V |
F |
Growth 25 °C microaerobic |
– |
– |
+ |
– |
– |
– |
– |
U |
– |
– |
+ |
+ |
+ |
+ |
– |
– |
– |
– |
F |
– |
Growth 37 °C microaerobic |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
V |
+ |
+ |
+ |
+ |
– |
+ |
+ |
– |
+ |
+ |
Growth 42 °C microaerobic |
+ |
+ |
+ |
+ |
+ |
+ |
M |
+ |
M |
V |
M |
V |
– |
– |
M |
+ |
+ |
– |
+ |
+ |
Growth 37 °C anaerobic |
+ |
+ |
+ |
– |
+ |
– |
+ |
+ |
– |
+ |
F |
+ |
M |
+ |
+ |
– |
– |
+ |
– |
w |
Growth 37 °C aerobic |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
V |
– |
– |
– |
– |
– |
– |
– |
– |
Growth on nutrient agar |
+ |
U |
U |
– |
– |
+ |
F |
w |
+ |
+ |
M |
U |
M |
U |
+ |
M |
+ |
U |
+ |
+ |
Growth on MacConkey |
– |
U |
+ |
– |
+ |
V |
– |
– |
– |
M |
M |
M |
V |
– |
M |
– |
– |
– |
V |
V |
R to nalidixic acid (30 µg) |
-* |
– |
– |
– |
V |
– |
M |
+ |
V |
+ |
+ |
+ |
V |
+ |
V |
– |
V |
V |
+ |
+ |
R to cephalothin (30 µg) |
+ |
R |
– |
+ |
– |
+ |
– |
– |
M |
– |
– |
U |
– |
– |
– |
– |
M |
– |
F |
– |
H2 requirement |
– |
– |
– |
V |
– |
– |
+ |
– |
– |
+ |
– |
– |
– |
– |
+ |
– |
– |
+ |
– |
U |
DNA G+C content (mol%) |
27.4 |
28 |
29 |
35 |
U |
31 |
37–41 |
31.9 |
32.4 |
45–46 |
33–35 |
33 |
33–34 |
33.6 |
44–46 |
34 |
27.9 |
32–33 |
34–36 |
31–33 |
Selenite reduction |
– |
V |
U |
– |
U |
+ |
F |
U |
– |
– |
M |
U |
F |
– |
– |
– |
U |
– |
+ |
+ |
Growth on 2 % NaCl |
+ |
– |
U |
– |
– |
– |
F |
+ |
– |
V |
– |
U |
F |
+ |
– |
F |
– |
U |
– |
– |
Growth on 1 % glycine |
+ |
+ |
w |
– |
V |
M |
F |
+ |
– |
+ |
+ |
+ |
F |
+ |
+ |
V |
+ |
+ |
+ |
F |
Growth on 0.04 % TTC |
F |
V |
U |
+ |
U |
+ |
– |
– |
V |
+ |
– |
+ |
– |
– |
– |
– |
+ |
– |
F |
– |
TTC reduction |
F |
V |
U |
– |
U |
+ |
– |
– |
V |
M |
– |
+ |
– |
– |
– |
– |
U |
– |
F |
– |
Characteristic |
21 |
22 |
23 |
24 |
25 |
26 |
27 |
28 |
29 |
30 |
31 |
32 |
33 |
34 |
35 |
36 |
37 |
38 |
39 |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
Motility |
U |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
U |
U |
U |
U |
+ |
+ |
+ |
+ |
+ |
– |
+ |
Oxidase |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
– |
+ |
+ |
+ |
+ |
V |
+ |
+ |
+ |
+ |
+ |
Catalase |
+ |
+ |
M |
+ |
+ |
+ |
+ |
– |
+ |
+ |
– |
+ |
F |
+ |
V† |
+ |
– |
F |
+ |
Urease |
– |
– |
– |
– |
– |
– |
– |
– |
+ |
U |
+ |
+ |
– |
– |
V‡ |
U |
– |
+ |
U |
Nitrate reduction |
+ |
+ |
– |
+ |
+ |
+ |
+ |
F |
V |
U |
+ |
+ |
+ |
+ |
M |
U |
+ |
+ |
+ |
Hippurate hydrolysis |
– |
– |
+ |
+ |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
– |
Indoxyl acetate hydrolysis |
– |
– |
+ |
+ |
– |
U |
F |
– |
– |
U |
– |
– |
+ |
V |
– |
– |
+ |
V |
– |
Alkaline phosphatase |
U |
U |
– |
– |
+ |
U |
– |
– |
– |
U |
U |
U |
– |
– |
– |
U |
– |
– |
– |
H2S production TSI |
+ |
– |
– |
– |
– |
U |
– |
+ |
– |
U |
+ |
+ |
– |
V |
+ |
– |
– |
– |
– |
Alpha haemolysis |
+ |
nd |
+ |
+ |
+ |
U |
M |
F |
– |
U |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
V |
U |
Growth 25 °C microaerobic |
+ |
– |
– |
– |
– |
– |
– |
– |
– |
U |
+ |
+ |
– |
– |
– |
– |
– |
– |
– |
Growth 37 °C microaerobic |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
– |
V |
+ |
+ |
+ |
+ |
+ |
Growth 42 °C microaerobic |
– |
– |
– |
+ |
+ |
+ |
+ |
+ |
+ |
+ |
– |
– |
w |
V |
M |
+ |
M |
V |
+ |
Growth 37 °C anaerobic |
w |
– |
– |
– |
w |
U |
– |
+ |
+ |
U |
+ |
+ |
+ |
+ |
+ |
+ |
– |
+ |
+ |
Growth 37 °C aerobic |
– |
– |
– |
– |
– |
– |
– |
– |
– |
U |
– |
– |
– |
– |
– |
– |
– |
– |
– |
Growth on nutrient agar |
U |
+ |
+ |
+ |
U |
+ |
+ |
+ |
U |
+ |
U |
U |
F |
V |
M |
– |
+ |
M |
– |
Growth on MacConkey |
U |
U |
– |
– |
U |
U |
U |
M |
U |
U |
U |
U |
– |
+ |
V |
F |
– |
V |
w |
R to nalidixic acid (30 µg) |
+ |
+ |
– |
– |
+ |
– |
M |
M |
U |
M |
– |
– |
M |
– |
M |
+ |
– |
– |
+ |
R to cephalothin (30 µg) |
– |
+ |
– |
+ |
+ |
+ |
+ |
– |
U |
F |
– |
– |
– |
– |
– |
– |
F |
– |
+ |
H2 requirement |
– |
– |
– |
– |
U |
U |
– |
+ |
– |
U |
– |
– |
+ |
+ |
– |
U |
– |
+ |
– |
DNA G+C content (mol%) |
36 |
U |
28–30 |
30–31 |
36 |
30 |
31–33 |
38–39 |
29.5 |
29 |
31 |
30 |
45–46 |
44–46 |
30–33 |
30 |
32–35 |
28–30 |
29 |
Selenite reduction |
U |
U |
– |
M |
U |
U |
V |
F |
U |
U |
U |
U |
– |
– |
U |
– |
+ |
– |
+ |
Growth on 2 % NaCl |
U |
– |
– |
– |
– |
+ |
+ |
– |
U |
M |
U |
U |
M |
+ |
+ |
+ |
– |
+ |
– |
Growth on 1 % glycine |
+ |
+ |
F |
M |
– |
+ |
+ |
V |
+ |
+ |
– |
V |
M |
V |
+ |
M |
+ |
+ |
– |
Growth on 0.04 % TTC |
U |
U |
V |
M |
U |
U |
M |
– |
U |
U |
U |
U |
– |
– |
– |
U |
– |
+ |
– |
TTC reduction |
U |
U |
V |
M |
U |
U |
M |
– |
U |
U |
U |
U |
– |
– |
– |
U |
– |
– |
– |