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Table 1.

Typical nocturnal roosting sites of different butterfly species

Species Roosting substrate Sources
Hesperiidae
Celaenorrhinus fritzgaertneri in a small cave (DeVries et al. 1987)
Pyrgus malvae at the top of dead flower-heads (Hoskins 2018)
Erynnis tages at the top of dead flower-heads (Hoskins 2018)
Nymphalidae
Clossiana euphrosyne on bracken fronds or on the flowers of rushes (Hoskins 2018)
Coenonympha pamphilus in a head-downwards posture at the top of grass heads (Hoskins 2018)
Danaus plexippus gregariously on various types of trees or shrubs (with a general preference for maples and conifers, pecans and oaks) (Davis et al. 2012)
Heliconius charitonia gregariously on leafless twigs of Anguria trees (Waller and Gilbert 1982)
Heliconius erato gregariously on leafless fine twigs or tendrils of dead vines (Finkbeiner et al. 2012; Mallet 1986)
Heliconius ethilla gregariously on leafless twigs (Turner 1975)
Heliconius sara shaded areas with plenty of thin dry vines and branches under relatively dense vegetation mats (Salcedo 2010b)
Lasiommata megera under leaves, under the lower boughs of trees or crevices in banks and walls, on fences (Dennis 1986)
Manataria maculata in shaded embankments, tree holes, and other dark hiding places (Hanson 2000)
Maniola jurtina in a head-downwards posture at the top of grass heads (Hoskins 2018)
Marpesia berania on the underside of the leaves of rubiaceous trees or other small trees (Benson and Emmel 1973)
Melanargia galathea in a head-downwards posture at the top of grass heads (Hoskins 2018)
Smyrna karwinskii gregariously in cavities of lava walls, tree trunks and on the underside of concrete slabs roofing alleys between sheds (Muyshondt and Muyshondt 1974)
Papilionidae
Papilio polyxenes asterius primarily on inflorescences, or apices, but also on stems, scapes or culms of daisies, other herbs and grasses, roost singly (Rawlins and Lederhouse 1978)
Pieridae
Colias eurytherne singly or gregariously in dense grass, in dense crown vetch, on the leaves of emergent forbs (Clench 1970)
Phoobis sennae eubule gregariously on the yellow-green leaves of vines (Clench 1970)
Pieris rapae gregariously on leaves or stems of the upper branches of Pittosporurn undulatum Vent. Var. val’iegatum (McFarland 1971)
Lycaenidae
Lycaenidae icarus on the flower-heads and stems of grasses and other plants (Frohawk 1914)
Lysandra bellargus in small groups on tall vegetation (Thomas 1983)
Plebejus argus on shrubs (bramble, gorse) and tall herbs (rank bunched grasses such as Dactylis glomerata, bracken and flowering herbs) (Dennis 2004)
Pseudophilotes sinaicus on the tips of dead stalks/dry flower heads of Jasonia montana (James 2006)
Zizina otis on low vegetation or on bushes (Hoskins 2018)