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D-Glucuronic acid and D-N-acetylglucosamine are linked by β-1,3 bonds (blue) to form a disaccharide. Multiple disaccharides are linked by β-1,4 bonds (red) to form hyaluronic acid. Mammalian and microbial hyaluronidases cleave β-1,4 bonds (red), and leech/hookworm hyaluronidases degrade β-1,3 bonds (blue).
