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. 2020 Jun 29;10(7):1110. doi: 10.3390/ani10071110

Table 2.

Behavioral Definitions. Definitions follow those of prior research [20]. Behaviors were coded as duration (D) or frequency (F).

Code Behavior Behavioral Definition
EX Exploration (D) Motor activity directed toward physical aspects of the environment, including sniffing, and gentle oral examination, such as licking
LO Locomotion (D) Walking or running around without exploring the environment
OB Oral behavior (D) Any vigorous behavior directed toward the environment/cage using the mouth (including chewing, biting, shaking, pulling with the mouth)
SC Scratching (D) All active behaviors resulting in physical contact with the cage/door, including scratching the cage/door with the paws, jumping on the cage/door, handling with the forelimbs
OE Oriented to the environment (D) Sitting, standing, or lying down (the head does not rest on the ground) with obvious orientation toward the physical or social environment, including sniffing, close visual inspection, distant visual inspection (vigilance or scanning)
PA Passive behavior (D) Lying down with the head on the ground without any obvious orientation toward the physical or social environment
PT Panting (D) Panting
WH Whining (D) Whining
BA Barking (D) Barking
CI Circling (D) Movement of the dog in circles
PL Play (D) Any vigorous or galloping gaited behavior directed towards a toy; including chewing, biting, shaking from side to side, scratching or batting with the paw, chasing rolling balls, and tossing using the mouth. Although the dog may take the objects into its mouth, destruction is not included in this category
GR Grooming (D) Action of cleaning the body surface by licking, nibbling, picking, rubbing, scratching, et cetera, directed toward the animal’s body
Ears back (D) Ears flattened and back
HO Howling (D) Howling
TR Trembling (D) Trembling/shaking movements of the body or head
PU Paw up (D) Front limb raised
LL Lip licking (F) Part of tongue is shown and moved along the upper lip
YA Yawning (F) Yawning
EL Elimination (F) Defecation or urination in sitting or standing position