Table 2:

Behaviors recorded and their definitions

Behavior	Definition
Aggression
Erh erhb	The animal emits low-pitched, staccato chattering (Epple, 1968; Bezerra & Souto, 2008; Stevenson & Poole, 1976).
Aggressionb	The animal lunges at (Abbott, 1984), snap-bites (Stevenson & Poole, 1976), chases (Saltzman et al, 1997) cuffs (Stevenson & Poole, 1976) or fights (Abbott, 1984) another animal.
Individual
Autogrooma	The animal cleans its own fur or skin with hand or mouth (including scratching- rapidly drawing claws of hand or foot across the fur or skin).
Inactivea	The animal remains stationary while alone (either resting, with tail curled around the body, or vigilant to surroundings- including the observer), without engaging in any other behavior (Saltzman et al, 1997).
Locomotiona	The animal travels between locations by walking, running, climbing or jumping.
Foragea	The animal is engaged in any activity directly related to acquiring or ingesting food.
Solitary playa	High activity behavior performed alone, such as hanging or swinging on a rope, chasing tail (Stevenson & Poole, 1976), or investigating objects in the environment by handling, sniffing, gnawing or attending to them whilst walking around them.
Othera	The animal has entered the nestbox and so cannot be seen by the observer, or is performing any other behavior not noted here.
Sexual
Mountb	The animal climbs/attempts to climb onto another animals back from behind and grips the other animal around the waist, may include pelvic thrusting (Kendrick & Dixson, 1983).
Social affiliative
Calm vocalb	The animal emits quiet, birdlike calls, such as ‘trills’ and ‘chirps’ audible to the observer.
Proximitya	The animal is stationary, sitting, crouching or lying next to another individual, with some form of physical contact (often torso-torso) for at least 3 secs (huddle- Abbott, 1984).
Allogrooma	The animal cleans the fur or skin of another individual with its hands or mouth (Abbott, 1984).
Social playa	High activity social interactions involving close, non-aggressive physical contact with other individuals, such as wrestling, chasing, grasping, pouncing, back-hugging, batting, biting or mutual exploration, often accompanied with a play face (open mouth without lip retraction) (Stevenson & Poole, 1976).
Submission
Nga ngab	The animal emits a relatively low-pitched, infantile squeal (Epple, 1968).
Submitb	The animal moves at least one body length away from another animal within 1 second of the other animal being in proximity (Saltzman et al, 1997). May include the animal flattening their tufts against their head, partially opening their mouth with corners retracted, exposing teeth (facial grimace) or half closing their eyelids (eye slit) (Stevenson & Poole, 1976; Abbott, 1984).

^*a.duration used in analysis;

^b.frequency used in analysis