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. 2020 Jan 27;69(5):1016–1032. doi: 10.1093/sysbio/syaa002

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© The Author(s) 2020. Published by Oxford University Press, on behalf of the Society of Systematic Biologists.

This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted reuse, distribution, and reproduction in any medium, provided the original work is properly cited.

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Figure 1. — The basic logic of the SPR proposal mechanisms. a) First, a branch (with length ) is picked at random from all branches in the tree. One of the subtrees incident to is then picked at random and pruned away; this subtree is labeled A here. b) For regrafting, a branch is picked in the other subtree (labeled BCDE). The branch in the moving direction (the pendant branch) is moved together with , and c) inserted adjacent to , resulting in connecting and . The specific SPR variants differ in how they pick .

Inline graphic — The basic logic of the SPR proposal mechanisms. a) First, a branch (with length ) is picked at random from all branches in the tree. One of the subtrees incident to is then picked at random and pruned away; this subtree is labeled A here. b) For regrafting, a branch is picked in the other subtree (labeled BCDE). The branch in the moving direction (the pendant branch) is moved together with , and c) inserted adjacent to , resulting in connecting and . The specific SPR variants differ in how they pick .