Eco-evolutionary dynamics of nested Darwinian populations and the emergence of community-level heredity

. 2020 Jul 7;9:e53433. doi: 10.7554/eLife.53433

Each particle of type i . . .	With rate. . .
Reproduces (add a particle of type i) Dies (remove a particle of type i)	$r_{i}$ $r_{i} (\sum_{j} δ_{c_{i} = c_{j}} x_{j} a_{j}^{intra} K^{- 1} + \sum_{j} δ_{c_{i} \neq c_{j}} x_{j} a_{j}^{inter} K^{- 1})$