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. 2020 Sep 21;10:15402. doi: 10.1038/s41598-020-72373-0

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Anions located along the TSP2 threefold axis support trimer oligomerization. (a) A chloride located between the neck and the D3′ domain forms charge-charge interactions with three guanidinium groups of Arg284. (b) A chloride located along the intermolecular D3 β-helices axis interacts with three side chain amide groups of Asn565. (c) A sulfate located along the intermolecular D3 β-helices axis form charge-charge interaction with the carboxyl groups of Asp415 and the amino groups of Lys465. The figure was generated using the computer program PyMol v1.8.0.2 (Schrödinger, LLC, https://www.pymol.org).