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. 2020 Sep 23;10:329. doi: 10.1038/s41398-020-01012-z

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 4 — a F2 social novelty testing revealed that IUO offspring spent less time with naive animals and more time with the cagemate than the other groups. IUO also had fewer contacts with the naïve animal. b F2 social preference testing showed IUO spent less time with the naive animal than the other groups, and IUO had more contacts with the toy than did the other groups. *p < 0.05; **p < 0.01; ***p < 0.001 as determined by two-way ANOVA followed by a post hoc Tukey’s test. c Marble burying tests in both generations showed increased burying activity in the PNO and IUO groups. **p < 0.01; ***p < 0.001; ****p < 0.0001 as determined by two-way ANOVA followed by a post hoc Tukey’s test.