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. 2020 Sep 30;11:4905. doi: 10.1038/s41467-020-18707-y

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 5 — a Initially, Brf1 and TBP are recruited to the N-terminal TPR array of τ131. The distance between the τ95 DBD and the τ131 TPR array serves as a molecular ruler that places TFIIIB within a certain distance upstream of the TSS. b TBP binds and bends the upstream DNA sequence. The lifetime of this complex depends on the upstream DNA sequence engaged. c Bdp1 then enters the complex and stabilizes the bent state. d Recruitment of Pol III and promoter opening displaces τA, freeing the transcribed region.