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. 2020 Sep 13;8(3):19. doi: 10.3390/jdb8030019

Table 1.

Known agonists and antagonists of BMP-2.

Protein/Gene Type of Protein Function References
Brorin Glycoprotein Inhibits BMP-2 activity [184,185,186]
Cerberus Cytokine Blocks BMP-2 signaling [187,188]
Chordin Glycoprotein Binds directly to BMP-2 to prevent its activity [170,171,177,189,190]
DAN Family Glycoproteins Binds directly to BMP-2 and inhibits its activity [191]
Follistatin Glycoprotein Binds directly to BMP-2 to prevent its activity [173,192,193,194]
FLRG Glycoprotein Downregulates BMP-2 and limits its activity [194,195,196]
Grem2 Glycoprotein Inhibits BMP-2 activity [197,198]
Noggin Glycoprotein Binds directly to BMP-2 to prevent its activity [9,172,199,200]
Sclerostin (SOST) Glycoprotein Binds to LRP5/6 to prevent Wnt and BMP-2 signaling [201,202,203,204]
Shrew Shrew-1: Transmembrane protein Enhances peak BMP-2 signaling activity [169,170]
Twisted Gastrulation Produces small cysteine rich protein Can enhance or inhibit BMP-2 activity [169,205,206,207,208,209,210]
USAG-1 Glycoprotein Physically interacts with BMP-2 to prevent signaling [211,212]