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(A) RDA plot based on the relative abundances of all ammonia-oxidizers (qPCR results); (B) CCA plot based on comammox Nitrospira clade A OTU abundances (high-throughput sequencing results). Black circles: samples, Blue arrows: ammonia-oxidizers, Blue triangles: Comammox Nitrospira clade A OTUs, Red arrows: environmental factors
