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. 2020 Oct 19;10:17607. doi: 10.1038/s41598-020-74551-6

Table 1.

Annotated checklist of known species of Liadopsyllidae Martynov, 19265.

Taxon Locality Age and formation Preservation Sex
Amecephala gen. nov
A. pusilla sp. nov. Myanmar, Kachin State, Hukawng Valley, SW of Maingkhwan, Noije Bum 2001 Summit Site amber mine Mid-Cretaceous (Aptian/Cenomanian) Amber inclusion Female
Gracilinervia Becker-Migdisova, 19857, p. 78
G. mastigimatoides Becker-Migdisova, 19857, p.s 79 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Unknown
Liadopsylla Handlirsch, 19216, p. 213
L. apedetica Ouvrard, Burckhardt et Azar, 2010 in Ouvrard et al. 20108, p. 173 Lebanon, Mdeyrij-Hammana, Casa Baabda Lower Cretaceous (late Barremian); Grès du Liban Fm Amber inclusion Female
L. asiatica Becker-Migdisova, 19857, p. 74 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Unknown
L. brevifurcata Becker-Migdisova, 19857, p. 69 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Impression Unknown
L. geinitzi Handlirsch, 19216, p. 213 Germany, Mecklenburg, Dobbertin Lower Jurassic (early Toarcian); Harpoceras falciferum ammonoid zone Impression Unknown
L. grandis Becker-Migdisova, 19857, p. 63 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Male
L. hesperia Ouvrard et Burckhardt, 2010 in Ouvrard et al. 20108, p. 175 U.S.A., New Jersey, Middlesex County, Sayreville Upper Cretaceous (Turonian); Raritan Fm Amber inclusion Female?
L. karatavica Becker-Migdisova, 19857, p. 73 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Male
L. lautereri (Shcherbakov, 2020) 9, p. 130 Russia, SW Buryatia Zakamensk district, Khasurty, 10 km S of Tsakir Lower Cretaceous (Aptian); Gusinoe Ozero Gr Impression Unknown
L. loginovae (Shcherbakov, 2020) 9, p. 132 Russia, SW Buryatia Zakamensk district, Khasurty, 10 km S of Tsakir Lower Cretaceous (Aptian); Gusinoe Ozero Gr Impression Unknown
L. longiforceps (Becker-Migdisova, 1985)7, p. 61 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Unknown
L. mongolica Shcherbakov, 198813, p. 61 Mongolia, Bon Tsagaan Lower Cretaceous (Aptian); Dzun-Bain Fm., Khurilt Mb Impression Unknown
L. obtusa Ansorge, 199614, p. 55 Germany, Mecklenburg-Vorpommern, Grimmen, Klein Lehmhagen pit Lower Jurassic (early Toarcian); Dactylioceras tenuicostatum zone Impression Unknown
L. tenuicornis Martynov, 19265, p. 1359 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Male
L. turkestanica Becker-Migdisova, 194915, p. 42 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression 4 Males, 4 females
Malmopsylla Becker-Migdisova, 19857, p. 76
M. karatavica Becker-Migdisova, 19857: 76 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Impression Unknown (forewing only)
Mirala Burckhardt et Poinar, 20194, p. 3
M. burmanica Burckhardt et Poinar, 20194, p. 3 Myanmar, Kachin State, Hukawng Valley, SW of Maingkhwan, Noije Bum 2001 Summit Site amber mine Mid-Cretaceous (Aptian/Cenomanian) Amber inclusion Unknown
Neopsylloides Becker-Migdisova, 19857, p. 77
N. turutanovae Becker-Migdisova, 19857, p. 77 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Unknown
Pauropsylloides Becker-Migdisova, 19857, p. 79
P. jurassica Becker-Migdisova, 19857, p. 79 Kazakhstan, Karatau, Kasharata (Mikhailovka), Aulie Upper Jurassic (Callovian); Karabastau Fm Compression Unknown
Stigmapsylla Shcherbakov, 20209, p. 129
S. klimaszewskii Shcherbakov, 20209: 130 Russia, SW Buryatia Zakamensk district, Khasurty, 10 km S of Tsakir Lower Cretaceous (Aptian); Gusinoe Ozero Gr Impression Unknown

All known specimens are adults. Becker-Migdisova (1985, p. 62)7 synonymised Asientomum Martynov, 19265, p. 1364 (replacement name for Lithentomum Martynov, 19265, p. 1365, nec Scudder, 186716, p. 206) with Liadopsylla and transferred Lithentomum praecox Martynov, 19265, p. 1365, to Liadopsylla. The venation of the forewing and particularly of the hindwing as described by Martynov5 clearly places this species in the Psocodea. Pending an examination of the holotype we follow Martynov5 rather than Becker-Migdisova7 in this matter.