Table 4.

The functions of candidate genes identified in this study.

QTLs	Gene	Position	Function	Reference
1	CHST111	54433922-54593321	Catalyze the transfer of sulfate to position 4 of the GalNAc residue of chondroitin.	Zhou et al., 2016
2	DSG4	106052985-106078954	Haplotypes related with the color of the goat coat.	E et al., 2016
2	CABLES11	102881201-102953764	An adapter protein that links cyclin-dependent kinase with nonreceptor tyrosine kinases and regulates the activity of Cdks by enhancing their Y15 phosphorylation.	Huang et al., 2017
2	GATA6	102636810-102654409	Promotes hair follicle progenitor cell renewal.	Wang et al., 2017
2	GREB1L	102280644-102409696	A coactivator for retinoic acid receptors.	Brophy et al., 2017
2	YES1	101881440-101930073	Mediates the biological process of tyrosine.	Patel et al., 2013, Ta et al., 2018
20	EDN3	10799441-10816861	Variation in EDN3 resulted in dermal hyperpigmentation phenotype or fibromelanosis	Dorshorst et al., 2011, Lillie et al., 2017, Shinomiya et al., 2012
20	BCL2L11	10015061-10033228	Homologous to BCL2 gene related with gray hair in mouse	Seiberg et al., 2013
20	CDH221	10647698-10705460	The expression was associated with melanoma metastasis	Piche et al., 2011

¹Genes overlapped with the significant F_ST clusters.