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. 2020 Sep 29;7(10):63. doi: 10.3390/medicines7100063

Table 2.

Flavones isolated from Stachys spp.

Species Plant Parts Compound Ref
Subgenus Stachys
Section Ambleia
S. aegyptiaca Pers. Aerial parts Apigenin (1),
Apigenin 7-O-β-D-glucoside (cosmoside) (2),
Apigenin 7-O-[6′″-O-acetyl]-allosyl-(1→2)-β-D-glucoside (3),
Apigenin 6,8-di-C-glucoside (Vicenin-2) (10),
Isoscutellarein 7-O-allosyl-(1→2)-β-D-glucoside (13),
Isoscutellarein-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucoside (15),
Luteolin (34),
Luteolin-7-O-[6′″-O-acetyl]-allosyl]-(1→2)-β-D-glucoside (39),
6,8 Di-C-β-D-glucopyranosyl luteolin (Lucenin-2) (40),
Chrysoeriol (42)
Chrysoeriol 7-O-β-D-glucoside (43),
Hypolaetin 7-O-[6′″-O-acetyl]-allosyl-(1→2)-[3″-O-acetyl]- β-D-glucoside (54),
Apigenin 7-O-diglucoside (not determined),
Luteolin 7-O-diglucoside (not determined)
[68]
Aerial parts Apigenin-7-(3″-E-p-coumaroyl)-β-D-glucoside (4),
Apigenin 7-(6″-p-coumaroyl)-β-D-glucoside (6)
[69]
Aerial parts Isoscutellarein (11),
3′,4′-Dimethyl-luteolin-7-O-β-D-glucoside (41)
[70]
Isoscutellarein 8-O-(6″-trans-p-coumaroyl)-β-D-glucoside (18) [71]
S. inflata Beth. Scutellarein 7-O-β-D-mannopyranosyl-(1→2)-β-D-glucoside (stachyflaside) (31) [72]
Isoscutellarein (11), 4′-Μethyl-isoscutellarein (12), Scutellarein (29) [73]
S. schtschegleevii Sosn. ex Grossh. Stems Apigenin 7-O-β-D-glucoside (2),
Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6),
3′-Hydroxy-isoscutellarein-7-O-[6′″-O-acetyl]-β-D-glucopyranoside (14),
Chrysoeriol 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (47)
[74]
Section Campanistrum
S. arvensis (L.) L. Aerial parts # 8-Hydroxyflavone-allosylglucosides (not determined) [75]
S. ocymastrum (L.) Briq. (= S. hirta L.) Aerial parts # 8-Hydroxyflavone-allosylglucosides (not determined) [75]
Aerial parts Apigenin (1),
Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6),
Isoscutellarein 7-O-allosyl-(1→2)- glucopyranoside (13),
Luteolin (34)
[76]
Section Candida
S. candida Bory & Chaubard Aerial parts Chrysoeriol (42), Chrysoeriol 7-(3″-E-p-coumaroyl)-β-D-glucopyranoside (46) [77]
Aerial parts Apigenin 7-O-β-D- glucopyranoside (2),
Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
Isoscutellarein 7-O-[6′″-O-acetyl]-allosyl-(1→2)-[6″-O-acetyl]- glucopyranoside (17),
4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21),
Chrysoeriol 7-O-β-D- glucopyranoside (43),
Chrysoeriol 7-(3″-E-p-coumaroyl)-β-D-glucopyranoside (46),
4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56)
[78]
S. chrysantha Boiss. and Heldr. Aerial parts Isoscutellarein 7-O-[6′″-O-acetyl]-allosyl(1→2)-[6″-O-acetyl]-glucoside (17), Luteolin 7-O-β-D-glucoside (37), Chrysoeriol (42), Chrysoeriol 7-O-β-D- glucopyranoside (43), Chrysoeriol 7-(3″-E-p-coumaroyl)-β-D-glucopyranoside (46) [77]
S. iva Griseb. Flowering aerial parts Apigenin (1), Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-[6″-O-acetyl]-β-D-glucopyranoside (17),
4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21),
4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56)
[56]
Section Corsica
S. corsica Pers. Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21)
[79]
Section Eriostomum
S. alpina L. Aerial parts # 8-Hydroxyflavone-allosylglucosides (not determined) [75]
Leaves # Hypolaetin 7-O-acetyl-allosyl-(1→2)-glucoside (not determined), Isoscutellarein-7-O-acetyl-allosyl-glucoside (not determined), Hypolaetin-4′-methyl- 7-O- acetyl-allosyl-glucoside (not determined) [5]
S. byzantina K. Koch. Aerial parts Apigenin (1), Apigenin 7-O-β-glucoside (2), Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6) [33]
Aerial parts Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6),
Isoscutellarein 7-O-β-D-allopyranosyl-(1→2)-[6″-O-acetyl]-β-D-glucopyranoside (16),
4′-Methyl-isoscutellarein-7-O-β-D-allopyranosyl-(1→2)-[6″-O-acetyl]-β-D-glucopyranoside (20)
[80]
S. cretica subsp. smyrnaea Rech. f. Aerial parts # Apigenin (1) [81]
S. germanica L. Aerial parts # Hypolaetin 7-allosyl-(1→2)-glucoside monoacetyl, Isoscutellarein 7-allosyl-(1→2)-glucoside monoacetyl, Hypolaetin 7-allosyl-(1→2)-glucoside diacetyl, Isoscutellarein-7-allosyl-(1→2)-glucoside diacetyl(not determined) [75]
Leaves # Apigenin 7-O-glucoside (2), Chrysoeriol 7-O-acetyl-allosyl-glucoside (not determined), 4′-Methyl-hypolaetin 7-O-acetyl-allosyl-(1→2)-glucoside (not determined), Apigenin 7-O-p-coumaroyl-glucoside (not determined) [5]
S. heraclea All. Aerial parts # 8-Hydroxyflavone-allosylglucosides (not determined) [75]
S. lanata Crantz. (=S. germanica L. subsp. germanica) Aerial parts Apigenin 7-O-β-D-glucopyranoside (2), Apigenin 7-(3″-Z-p-coumaroyl)-β-D-glucopyranoside (5), Apigenin 7-(6″-Z-p-coumaroyl)-β-D-glucopyranoside (7),
Apigenin 7-O-(3′′,6′′-di-O-E-p-coumaroyl)-β-D-glucopyranoside (Anisofolin A) (8),
Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
Isoscutellarein 4′-methyl ether 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21), 4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56)
[82]
S. spectabilis Choisy ex DC. Epigeal parts Isostachyflaside (25),
Spectabiflaside (28), Scutellarein 7-O-β-D-mannopyranosyl-(1→2)-β-D-glucopyranoside (stachyflaside) (31)
[83]
S. thirkei K. Koch. Whole plant # Apigenin (1) [84]
S. tmolea Boiss. Aerial parts # Apigenin (1), Apigenin-7-O-glucoside (2) [85]
S. tymphaea Hausskn. (=S. germanica subsp. tymphaea (Hausskn.) R. Bhattacharjee) Flowering aerial parts Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21),
4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]- allopyranosyl -(1→2)-[6″-O-acetyl]-glucopyranoside (58)
[86]
Section Fragilicaulis
S. subnuda Montbret & Aucher ex Benth Aerial parts Ιsoscutellarein 7-O-allosyl-(1→2)-glucoside # (13), Isoscutellarein 7-O-[6″′-O-acetyl]-allosyl-(1→2)-glucoside (15), 4′-Methyl-isoscutellarein-7-O-β-D-allopyranosyl-(1→2)-β-D-glucoside # (19), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucoside (21), 4′-Methyl-isoscutellarein-7-O-[6′″-O-acetyl]-allosyl(1→2)-[6″-O-acetyl]-glucoside # (24) [87]
Section Olisia
S. atherocalyx C. Koch Stachyflaside (31) [72]
Diacetylstachyflaside (not determined), Diacetylspectabiflaside (not determined), Spectabiflaside (28) [88]
5,8,4′-Trihydroxy-3′-methoxy-7-O-(β-D-glucopyranosyl-2″-O-β-D-mannopyranosyl)-flavone (Spectabiflaside) (28), Acetyl-sectabiflaside (not determined), [89]
Acetyl-isostachyflaside (26), Di-acetyl-isostachyflaside (27),
Spectabiflaside (28)
[90]
Leaves # Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21),
4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56)
[91]
S. angustifolia M. Bieb. Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21) [92]
S. annua (L.) L. Epigeal parts 4′-Methyl-isoscutellarein (12), 7-O-β-D-glucopyranosyl-
5,6-dihydroxy-4′-methoxyflavone (Stachannin A) (32),
4′-Methoxy-scutellarein-7-[O-β-D-mannopyranosyl-(1→2)-β-D-glucopyranoside] (Stachannoside B) (33)
[93]
Leaves # Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
4′-Methyl-isoscutellarein-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21),
4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56)
[92]
Aerial parts 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21) [94]
Aerial parts 4′-O-Methyl-isoscutellarein-7-O-[4′″-O-acetyl]allopyranosyl-(1→2)-
glucopyranoside (Annuoside) (23)
[95]
Subterranean organs 4′-O-Methyl-isoscutellarein (12), 4′-O-Methyl-isoscutellarein
7-O-(6′″-O-acetyl)allopyranosyl-(1→2)-glucopyranoside (21)
[95]
S.annua (L.) L. subsp. annua Flowering aerial parts 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21), Hypolaetin 7-O-[6′″-O-acetyl]-allosyl-(1→2)-[6″-O-acetyl]-glucopyranoside (53),
4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56)
[52]
S. beckeana Dörfler & Hayek Leaves # Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (21), 4′-Methyl-hypolaetin-7-O-[6′″-O acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56) [92]
S. bombycina Boiss. Aerial parts Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6), Stachyspinoside (44) [96]
S. parolinii Vis. Leaves # Isoscutellarein 7-O-[6′″-O-acetyl]-β- D-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56) [92]
S. leucoglossa Griseb. Leaves # Isoscutellarein 7-O-[6′″-O-acetyl]-allosyl(1→2)-[6″-O-acetyl]-glucoside (17), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21), 4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56) [92]
S. neglecta Klok. ex Kossko (=S. annua (L.) L.) Apigenin (1),
Apigenin 7-O-β-D-glucoside (2), Luteolin (34), Luteolin 7-O-β-D-glucoside (37)
[97]
S. recta L. Leaves Isoscutellarein 7-O-[6′″-O-acetyl]-allosyl(1→2)-[6″-O-acetyl]-glucoside (17), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]- allosyl](1→2)-β-D-glucoside (21), 4′-Methyl-hypolaetin 7-O-[6′″-O-acetyl]-β-D-allosyl(1→2)-β-D-glucoside (56) [91,92]
Aerial parts Apigenin 7-(3″-E-p-coumaroyl)-β-D-glucopyranoside (4), Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6), Isoscutellarein 7-O-[allosyl(1→2)]-
glucopyranoside (13), Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allosyl-(1→2)-β-D-glucopyranoside (15), Isoscutellarein 7-O-[6′″-O-acetyl]-allosyl(1→2)-[6″-O-acetyl]- glucopyranoside (17),
4′-Methylisoscutellarein 7-
O-[allosyl-(1→2)]-
glucopyranoside (19),
4′-Methyl-isoscutellarein 7-O-β-D-[6″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (20), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]- allosyl-(1→2)-β-D-glucopyranoside (21), 4′-Methyl-isoscutellarein 7-O-[6′″-O-acetyl]-allosyl-(1→2)-[6″-O-acetyl]-glucoside (24), Hypolaetin 7-O-allosyl-(1→2)-glucopyranoside # (50), 4′-Methyl-hypolaetin 7-O-allosyl(1→2)-glucoside # (55), 4′-Methyl-hypolaetin-7-O-[6″-O-acetyl]-allosyl-(1→2)- glucopyranoside (57),
4′-Methyl-hypolaetin 7-O-[6′″-O-acetyl]-allosyl-(1→2)-[6″-O-acetyl]- glucopyranoside (58)
[14]
S. labiosa Bertol. (=S. recta subsp. labiosa (Bertol.) Briq.) Leaves Isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21), 4′-Methyl-hypolaetin-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (56) [92]
S. subcrenata Vis.(=S. recta L. subsp. subcrenata (Vis.) Briq.) Leaves Isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21), 4′-Methyl-hypolaetin-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (56) [92]
S. baldaccii (Maly) Hand.—Mazz. (=S. recta L. subsp. baldaccii (K. Maly) Hayek) Leaves # Isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl]-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein-7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21) [92]
S. spinosa L. Aerial parts Chrysoeriol 7-O-[6′″-O-acetyl-allosyl]-(1→2)-glucoside (Stachyspinoside) (44) [98]
Aerial parts Chrysoeriol 7-O-[6′′-O-acetyl-allosyl]-(1→2)-glucoside (Isostachyspinoside) (45) [99]
S. tetragona Boiss. & Hayek Leaves # Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-allopyranosyl-(1→2)-β-D-glucopyranoside (21) [92]
Aerial parts Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
Isoscutellarein 7-O-[6′″-O-acetyl]- β-D-allosyl-(1→2)-[6″-O-acetyl]-β-D glucopyranoside (17)
[100]
Section Swainsoniana
S. anisochila Vis. & Pancic Leaves Isoscutellarein 7-O-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (13),
Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15),
Isoscutellarein 7-O-[6′″-O-acetyl]- β-D-allosyl-(1→2)-[6″-O-acetyl]-β-D-glucopyranoside (17), 4′-Methyl-isoscutellarein-7-O-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (19),
Hypolaetin 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (51),
Hypolaetin 7-O-[6′″-O-acetyl]- β-D-allopyranosyl-(1→2)-[6″-O-acetyl]-β-D glucopyranoside (53), 4′-Methyl-hypolaetin-7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (56),
4′-Methyl-hypolaetin 7-O-[6′″-O-acetyl]-β-D-allosyl-(1→2)-[6″-O-acetyl]-β-D glucopyranoside (58)
[101]
Leaves Apigenin 7-O-(p-coumaroyl)-β-D-glucopyranoside (not determined) [5]
S. decumbens Pers. (=S. mollissima Willd.) Aerial parts # 8-Hydroxyflavone-allosylglucosides (not determined) [75]
S. menthifolia Vis. (=S. grandiflora Host.) Leaves # Isoscutellarein 7-O-β-D-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein-7-O-β-D-[6′″-O-acetyl]-β-D allopyranosyl-(1→2)-β-D-glucopyranoside (21) [92]
S. swainsonii Benth. subsp. swainsonii Aerial parts Apigenin (1), Apigenin 7-O-β-D-glucopyranoside (2), Apigenin 7-O-β-D-glucoside (2), Luteolin 7-O-β-D-glucopyranoside (37), Chrysoeriol (42), Chrysoeriol 7-O-β-D-glucopyranoside (43), Stachyspinoside (44) [102]
S. swainsonii subsp. argolica (Boiss.) Phitos and Damboldt Aerial parts Apigenin (1), Luteolin 7-O-β-D-glucopyranoside (37), Chrysoeriol (42), Chrysoeriol-7-O-β-D-glucopyranoside (43), Chrysoeriol 7-(3″-E-p-coumaroyl)-β-D-glucopyranoside (46) [102]
S. swainsonii subsp. melangavica D. Persson Aerial parts Apigenin (1), Apigenin 7-O-β-D- glucopyranoside (2), Luteolin 7-O-β-D-glucopyranoside (37), Chrysoeriol-7-O-β-D-glucopyranoside (43), Stachyspinoside (44) [102]
S. swainsonii subsp. scyronica (Boiss.) Phitos and Damboldt Aerial parts Apigenin (1), Apigenin 7-O-β-D- glucopyranoside (2), Luteolin 7-O-β-D-glucopyranoside (37), Chrysoeriol-7-O-β-D-glucopyranoside (43), Stachyspinoside (44) [102]
S. ionica Halácsy Aerial parts Apigenin (1), Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6), Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15), 4′-Methyl-isoscutellarein 7-O-β-D-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (21) [20]
Section Stachys
S. sieboldii Miq. (=S. affinis Bunge) Aerial parts Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allosyl]-(1→2)-β-D-glucoside (15), 4′-Methyl-isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucoside (21) [20]
S. mialhesii Noé Aerial parts Apigenin 7-(6″-E-p-coumaroyl)-β-D-glucopyranoside (6), Isoscutellarein 7-O-[6′″-O-acetyl]-β-D-allopyranosyl-(1→2)-β-D-glucopyranoside (15) [103]
S. palustris L. 5-(glycuroglucosyl)-7-methoxybaicalein (Palustrin) (63), 5-(glucuronosyl)-7-methoxybaicalein (Palustrinoside) (64) [104]
Leaves # Vicenin-2 (10), Apigenin 7-O-p-coumaroyl-β-D-glucopyranoside (not determined) [5]
S. sylvatica L. Aerial parts # 8-Hydroxyflavone-allosyl-glucosides (not determined) [75]
Leaves # Chrysoeriol 7-O-acetylallosylglucoside (not determined), Apigenin 7-O-p-coumaroyl-β-D-glucopyranoside (not determined) [5]
S. plumosa Griseb. Leaves # Apigenin-7-O-β-D-glucoside (2), Luteolin 7-O-β-D-glucoside (37), Chrysoeriol 7-O-acetyl-allosyl-glucoside (not determined), Isoscutellarein 7-O-acetyl-allosyl-glucoside (not determined), Apigenin 7-O-p-coumaroyl-β-D-glucopyranoside (not determined) [5]
Section Zietenia
S. lavandulifolia Vahl. Aerial parts Apigenin (1), Hydroxygenkwanin (Luteolin 7-Methyl ether) (35), Chrysoeriol (42) [13]
S. tibetica Vatke Roots Apigenin 7-O-β-D-glucoside (2) [66]
Subgenus Betonica
Section Betonica
S. alopecuros (L.) Benth. Aerial parts p-coumaroyl-glucosides (not determined) # [75]
Leaves # Isoscutellarein 7-O-glucoside (11a),
Luteolin 7-O-glucuronide (36),
Luteolin 7-O-glucoside (37), Chrysoeriol 7-O-glucoside (43),
Hypolaetin 7-O-glucoside (49),
Hypolaetin 7-O-glucuronide (49a),
Selgin 7-O-glucoside (59),
Tricin 7-O-glucuronide (60),
Tricin 7-O-glucoside (61),
Apigenin 7-O-p-coumaroyl glucopyranoside (not determined)
[5]
S. foliosa Regel. (=S. betoniciflora Rupr.; Betonica foliosa Rupr.) Four flavonoids (not determined) [105]
S. monieri (Gouan) P.W. Ball. (=S. officinalis (L.) Trevis subsp. officinalis) Aerial parts p-coumaroyl-glucosides (not determined) # [75]
S. officinalis (L.) Trevis (=Betonica officinalis L.) Apigenin (1),
5, 6, 4′-trihydroxyflavone-7-O-β-D-glucoside (30)
[20]
Leaves # Apigenin 8-C-glucoside
(Vitexin) (9), Luteolin 7-O-glucuronide (36),
Luteolin 6-C-glucoside (isoorientin) (38),
Tricin 7-O-glucuronide (60),
Tricin 7-O-glucoside (61),
Tricetin 3′,4′,5′-trimethyl-7-O-glucoside (62),
Apigenin 7-O-p-coumaroyl glucopyranoside (not determined)
[5]
Aerial parts p-coumaroyl-glucosides (not determined) # [75]
Section Macrostachya
S. scardica Griseb. (=Betonica scardica Griseb.) Leaves # Apigenin 8-C-glucoside (9), Luteolin 7-O-glucoside (37),
Luteolin 6-C-glucoside (38),
Hypolaetin 7-O-glucoside (49),
Selgin 7-O-glucoside (59),
Tricin 7-O-glucuronide (60),
Tricin 7-O-glucoside (61),
Tricetin 3′,4′,5′-trimethyl-7-O-glucoside (isolation) (62),
Apigenin 7-O-p-coumaroyl glucopyranoside (not determined)
[5]

# identified compounds by means of HPLC, LC-MS, etc.