(A) Alignment of the DSSDE motif in corin LDLR8 modules from selected vertebrate species. The DSSDE motif (red) in corin LDLR8 modules from selected vertebrate species (human, Hum; monkey, Mac; rat, Rat; mouse, Mou; pigeon, Pig; zebrafish, Zeb) is aligned. This motif corresponds to the DxSDE motif found in LDLR modules in many proteins. The Asp-Glu residues (indicated by black dots), corresponding to Asp685 and Glu686 in human corin, have been shown to be part of a Ca2+-binding cage in LDLR module structures of other proteins. (B and C) Corin WT and mutants were expressed in MDCK cells and analyzed by immunostaining and confocal microscopy. Quantitative data of relative corin expression on apical and basolateral membranes, indicated by the ratio of FBL/FTotal, were analyzed by Image J software. The data (mean ± SEM) were from five experiments. Statistical analysis was done with ANOVA. n.s., not significant vs. corin WT in respective groups.