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. 2020 Nov 2;10:18802. doi: 10.1038/s41598-020-75708-z

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© The Author(s) 2020

Open AccessThis article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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WSI score distribution for test, OOD-near and OOD-all sets compared to the actual bulk-measured values (lower values in gray). Scores are computed using the ensemble model for each trait (percent positive tiles by majority vote of the best three). Middle white lines reflect the median slide score. Whiskers extend to $1.5 \times I Q R$ . (a)–(c) breast test, OOD-near and OOD-all respectively; (d)–(f) lung test, OOD-near and OOD-all (respectively). In the test set, especially worth noting are miR-17-5p in both cohorts, MKI67, FOXA1, MYC, mir-29a-3p and ESR1 in breast as well as KRAS and CD274 (PD-L1) in lung. In the OOD sets most notable are MKI67, FOXA1, MYC, ESR1 and FOXC1 in breast as well as miR-17-5p and miR-21-5p in lung.