Table 1.

The variables of the MuSSCRat model and their interpretation.

Variable	Interpretation
	The continuous characters for one lineage
	The state of the continuous characters at time
	The number of continuous characters
	An matrix containing the continuous characters for the species in the tree
	The discrete character for one lineage
	The state of the discrete character at time
	An matrix containing the discrete character for the species in the tree
	The complete history of the discrete character (the state at the beginning and end of the process,
	and all of the changes in between) along the lineage
	The instantaneous-rate matrix of the discrete-character CTMC model
	The instantaneous rate of change from state to state
	The background rate of continuous-character evolution among all lineages
	The background rate of continuous-character evolution for lineage
	The relative rates of continuous-character evolution for each continuous character
	The relative rates of continuous-character evolution for each discrete state
	The evolutionary correlation matrix
	The evolutionary correlation between continuous characters and
	The discrete-state-independent evolutionary variance–covariance matrix