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The expanded regions of the selected hybrid structures RNA-BC3-α (A) and RNA- BC3-β (B) showing a realization of an ‘in-line attack’ conformation with Θ (O2′-P-O5′) ≥155°. The guanidinium groups of the two arginine residues are located close to the sugar-phosphate backbone of the bulged nucleotide C15 to form a guanidine-guanidinium dyad, essential for catalysis.
