Loss of mutualism in the forward (collapse) direction. Shown are the steady state species abundances of network A versus the bifurcation parameter κ as it increases from zero for six values of ζ, the amplitude of DN. Mutualism is lost for ζ > ζc, where 4.5 < ζc < 5.0. The network parameter values are , , γ = 1, h = 0.2, ρ = 0.5, μX = 10−4 and μY = 10−4. The initial conditions are randomly chosen from the basin of the high abundance steady state.